LOS ANGELES -- After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

A zoo visitor greets Gramma in 2015 at the San Diego Zoo when the tortoise was a spry 130-year-old. (Mark Ralston / AFP via Getty Images) Gramma, a Galápagos tortoise whose life at the San Diego Zoo ...